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Alpine ibex prefer high altitude meadows alternated
with steep rocky slopes (Couturier, 1962; Nievergelt 1966; Hoffman e Nievergelt,
1972; Grignolio et al., 1999). Males and females differ greatly in their
habitat choices (Pedrotti, 1995): females use space in a more constant
way and are more linked to cliffs, while males move more, selecting different
kinds of habitat according to the season and their phisiological needs
(Francisci et al., 1985). In spring adult males prefer the more plain
meadows (Nievergelt, 1966; Peracino et al., 1989; Pedrotti, 1995), and
during the summer, they significantly select alpine meadows (Mascellani,
1997; Grignolio et al., 1999).
Generally, all studies on habitat selection in Alpine ibex, underline
how there is a strong preference by this species for environments characterised
by rocky steep slopes, which are actively searched for by both sexes especially
during the winter season (Nievergelt, 1966; Schaller, 1977; Apollonio
& Grimod, 1984; Tosi et al., 1986). This preference may be explained
by the high sensibility of this species towards strong snow accumulations
(Pedrotti, 1995).
During the summer season, the average altitude
of the frequented areas is just below 3000 m a.s.l., with a range from
2300 to 3200-3300 m a.s.l. and, exceptionally, even more. In winter the
altitudinal range occupied by Alpine ibex goes from 1800 to 2500-3000
m a.s.l. (Couturier, 1962; Wiersema, 1983). The lowest altitudes are reached
in spring, season during which the minimal values are influenced by the
altitude of the bottom of the valley and by the presence of wood, as extended
forested areas are a limit difficult to cross for Alpine ibex (Nievergelt,
1966).
In areas with a high density of Alpine ibex, the altitude distribution
range during summer can be much larger and stable groups of Alpine ibex
can be found as low as at 2000 m a.s.l. (Bassano & Peracino, 1987).
The results of numerous studies on Alpine ibex populations recently established
following natural migration or reintroductions, appear to dismiss the
view of Alpine ibex not being apt to live at low altitudes (von Elsner-Schack,
1982; Choisy, 1983, 1986, 1990 e 1994; Terrier et al., 1994; Raye, 1994).
The presence and establishment of stable populations in areas with altitudinal
limits even lower than 1000 meters, suggests a high plasticity in habitat
selection by the species. It appears however that a necessary condition
is the presence of large sectors of steep cliffs, rich in niches, able
to create favourable microclimatic conditions (Choisy, 1994).
The subalpine, alpine, and subnival horizons with a mean altitude of 2000
meters, are however the most concerned with the presence of Alpine ibex
(Nievergelt, 1966; Pfeffer & Settimo, 1973; Hofmann & Nievergelt,
1972; Peracino et al., 1989; Francisci et al., 1985; Wiersema, 1983).
One of the most important factors influencing habitat selection by Alpine
ibex is indeed the gradient and the extension of the slopes. All the authors
emphasise that slopes have to be between 30 and 45 degrees, particularly
during the winter period. Furthermore, the slopes have to be interrupted
by porches and hides in order to favour environmental diversity (Couturier,
1962; Nievergelt, 1966; Hofmann & Nievergelt, 1972; von Elsner Schack,
1982; Wieserma, 1983 e 1984; Tosi et al., 1986; Pedrotti, 1995).
Slopes characterise the different areas chosen by the two sexes: females
use steep terrains during the whole year, while males frequent also plain
meadows, in accordance with their different seasonal energetic needs (Nievergelt,
1966; Peracino et al.,1989; Pedrotti, 1995).
Slope exposure, for most part of the year, does not influence greatly
habitat choice by Alpine ibex: all exposures, even if in a differing extent,
are used (Tosi et al., 1986). Detailed analyses of space use by Alpine
ibex, in different populations, suggest however that south exposed slopes
(south, south-east, south-west) are the most preferred during the whole
year and are the only used during winter (Nievergelt, 1966; Peracino et
al., 1989). The choice of the kind of exposure appears to be influenced
by orography and thus can vary locally (Pedrotti, 1995).
In most Alpine ibex populations, wood does not appear to be important
in habitat choice (Bachler, 1935; Rauch, 1937; Baumann, 1949; Kuster,
1961; Nievergelt, 1966; Pedrotti, 1995). Wood appears to be used by Alpine
ibex only when it is crossed by ditches or interrupted by cliffs (Couturier,
1962; Hoffmann e Nievergelt, 1972), especially in the case of larch woods
(Peracino et al., 1989; Bassano et al., 1992).
During winter seasons with scarce snowfalls, in areas with high density
populations, adult and old Alpine ibex males can be found in larch and
mixed larch-spruce (Abies alba) woods. Also in this case, however, the
presence of small cliffs if of primary importance in the habitat choice
(Bassano et al., 1992; Grignolio et al., 1999).
Some populations, more or less recently established in areas with low
altitudes and strong vegetal cover , however appear to behave differently.
For example, the Alpine ibex population of Hochlantschstock, in eastern
Austra, 30 km distance from Graz, lacking of suitable winter terrain,
uses coniferous and broadleaf forests for the whole winter, causing a
significant damage to the forestry renewal (Kofler, 1981). Recent studies,
based on the observation of marked Alpine ibex, have shown that in areas
hosting high density populations (Albris-Swiss National Park), Alpine
ibex uses forest for a good part of the year and particularly in the months
of May, June and September. In these periods the damages to the vegetation,
mostly due to Alpine ibex scratching and horning the plants, can be significant
(Abderhalden & Buchli, 1998).
Low snow-falls are an important factor for survival in most species of
the genus Capra (Schaller, 1977). Areas with precipitation higher than
1500 mm/year appear thus to be unfavourable for the establishment of Alpine
ibex populations, while the optimal precipitation range for Alpine ibex
goes from 100 to 1000 mm/year (von Elsner Schack, 1982). Villaret (1987),
did not find any significant correlation between precipitation and the
dynamics of Alpine ibex populations in France.
Intense precipitation can however have an effect on the survival of kids
during spring (Nievergelt, 1966). Furthermore, even if the correlation
between birth rate and precipitation appears not to be significant in
the Gran Paradiso population (rs=-0.33, p= 0.08), the lowest values of
birth rate are found in years characterised by heavy precipitation (Bassano
et al., 1992).
These contradictory results are almost certainly due to considering snow
and rain alike: snow precipitation are important not in terms of centimetres
of fallen snow but mostly in terms of height and persistence on the soil
of the snow cover (Choisy, 1990).
A significant correlation between mean height of snow cover and Alpine
ibex population dynamics in Gran Paradiso National Park has been shown
for males (rs= -0.52, p<0.05), females (rs= -0.69, p<0.05) and yearlings
(rs= - 0.67, p<0.05)(Bassano et al., 1992).
Places with heavy snow cover are avoided by Alpine ibex (Couturier, 1962;
Nievergelt, 1966; Hoffmann & Nievergelt, 1972). This avoidance appears
to be due to the alpine ibex being scarcely adapted to snow, as suggested
by its particular body mass/body surface ratio and the anatomy of its
limbs and hoves (Kofler, 1983).
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